It is indeed manifest that multitudes of species are related in the closest manner to other species that still exist, or have lately existed; and it will hardly be maintained that such species have been developed in an abrupt or sudden manner.

Nor should it be forgotten, when we look to the special parts of allied species, instead of to distinct species, that numerous and wonderfully fine gradations can be traced, connecting together widely different structures.

Many large groups of facts are intelligible only on the principle that species have been evolved by very small steps.

For instance, the fact that the species included in the larger genera are more closely related to each other, and present a greater number of varieties than do the species in the smaller genera.

The former are also grouped in little clusters, like varieties round species; and they present other analogies with varieties, as was shown in our second chapter.